Plants moderate to robust, weak-stemmed, yellow, yellowish brown to golden brown; capitulum varying from rounded, not 5-radiate and twisted to flat 5-radiate and straight branched. Stem pale green to pale brown; superficial cortex of weakly to moderately differentiated. Stem leaves triangular-lingulate, 0.9-1.3 mm; usually appressed; apex obtuse and often erose; hyaline cells efibrillose and nonseptate. Branches tapering or in more robust forms, frequently blunt, straight to arcuate, leaves slightly to moderately elongated at distal end. Branch fascicles with 2 spreading and 2 pendent branches. Branch stems green, with cortex enlarged with conspicuous retort cells. Branch leaves ovate to ovate-lanceolate; more than 1.8 mm; straight, stiff, not much undulate and reflexed to recurved; margins entire; hyaline cells on convex surface with a few end pores, but mostly numerous small to very small (often barely visible) pores or wall thinnings free from the commissures, on concave surface similar, but with pores generally fewer and larger; chlorophyllous cells triangular in transverse section, just reaching concave surface or slightly enclosed. Sexual condition dioicous. Spores 18-27 -m; both surfaces covered with rough, irregular verrucate plates of papillae, bifurcated Y-mark sculpture on distal surface; proximal laesura less than 0.5 spore radius.