With 1,839 species, Begonia is the fifth-largest angiosperm genus.The species are terrestrial (sometimes epiphytic) herbs or undershrubs, and occur in subtropical and tropical moist climates, in South and Central America, Africa, and southern Asia. Terrestrial species in the wild are commonly upright-stemmed, rhizomatous, or tuberous. The plants are monoecious, with unisexual male and female flowers occurring separately on the same plant; the male contains numerous stamens, and the female has a large inferior ovary and two to four branched or twisted stigmas. In most species, the fruit is a winged capsule containing numerous minute seeds, although baccate fruits are also known. The leaves, which are often large and variously marked or variegated, are usually asymmetric (unequal-sided).